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Lewis, S. M., and S. N. Austad. 1990. Sources of intraspecific variation in sperm precedence in red flour beetles. American Naturalist 135:351-359.
Lewis, S. M., and S. N. Austad. 1994. Sexual selection in flour beetles: the relationship between sperm precedence and male olfactory attractiveness. Behavioral Ecology 5:219-224.