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Barnes, A. I., and M. T. Siva Jothy. 2000. Density-dependent prophylaxis in the mealworm beetle Tenebrio molitor L (Coleoptera: Tenebrionidae): cuticular melanization is an indicator of investment in immunity. Proceedings Royal Society of London Series B Biological Sciences 267:177-182.
Drnevich, J. M., A. I. Barnes, and M. T. Siva Jothy. 2002. Immune investment and sperm competition in a beetle. Physiological Entomology 27:228-234.
Polis, G. A., J. D. Barnes, M. K. Seely, J. R. Henschel, and M. M. Enders. 1998. Predation as a major cost of reproduction in Namib Desert tenebrionid beetles. Ecology 79:2560-2566.