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Aloia, A., I. Colombini, M. Fallaci, and L. Chelazzi. 1999. Behavioural adaptations to zonal maintenance of five species of tenebrionids living along a Tyrrhenian sandy shore. Marine Biology 133:473-487.
Chelazzi, L., and I. Colombini. 1989. Zonation and activity patterns of two species of the genus Phaleria Latreille (Coleoptera; Tenebrionidae) inhabiting an equatorial and a Mediterranean sandy beach. Ethology Ecology and Evolution 1:313-321.
Chelazzi, L., G. Bertin, I. Colombini, and G. Poggesi. 1986. Researches on the coast of Somalia Heterocheira martellii Ardoin, 1974 (Coleoptera Tenebrionidae) First description of the larva and pupa. Monitore Zoologico Italiano Supplemento 21:1-10.
Chelazzi, L., I. Colombini, G. Poggesi, and G. Bertin. 1985. Researches on the coast of Somalia The shore and the dune of Sar Uanle 37 Tenebrionidae (Coleoptera. Monitore Zoologico Italiano Supplemento 20:191-210.
Colombini, I., A. Aloia, M. Fallaci, and L. Chelazzi. 1996. Spatial and temporal strategies in the surface activity of some sandy beach arthropods living along the French Atlantic coast. Marine Biology 127:247-257.
Colombini, I., L. Chelazzi, M. Fallaci, and L. Palesse. 1994. Zonation and surface activity of some tenebrionid beetles living on a mediterranean sandy beach. Journal of Arid Environments 28:215-230.
Colombini, I., L. Chelazzi, M. Fallaci, E. Lucarelli, and A. Mascagni. 1991. The Coleoptera inhabiting the beach-dune system of the Burano lagoon (GR): Population dynamics and zonation of the five most abundant families. Redia 74:87-110.
Colombini, I., M. Fallaci, and L. Chelazzi. 2005. Micro-scale distribution of some arthropods inhabiting a Mediterranean sandy beach in relation to environmental parameters. Acta Oecologica 28:249-265.
Fallaci, M., A. Aloia, I. Colombini, and L. Chelazzi. 2002. Population dynamics and life history of two Phaleria species (Coleoptera, Tenebrionidae) living on the Tyrrhenian sandy coast of central Italy. Acta Oecologica 23:69-79.
Fallaci, M., I. Colombini, L. Palesse, and L. Chelazzi. 1997. Spatial and temporal strategies in relation to environmental constraints of four tenebrionids inhabiting a Mediterranean coastal dune system. Journal of Arid Environments 37:45-64.