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Parmenter, R. R., C. A. Parmenter, and C. D. Cheney. 1989. Factors influencing microhabitat partitioning among coexisting species of arid-land darkling beetles (Tenebrionidae): behavioral responses to vegetation architecture. Southwestern Naturalist 34:319-329.
Parmenter, R. R., C. A. Parmenter, and C. D. Cheney. 1989. Factors influencing microhabitat partitioning in arid-land darkling beetles (Tenebrionidae): temperature and water conservation. Journal of Arid Environments 17:57-67.