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Edmunds, J., J. M. Cushing, R. F. Costantino, S. M. Henson, B. Dennis, and R. A. Desharnais. 2003. Park's Tribolium competition experiments: A non-equilibrium species coexistence hypothesis. Journal of Animal Ecology 72:703-712.
Edmunds, M. 2000. Why are there good mimics and poor mimics?. Biological Journal of the Linnean Society 70:459-466.