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Wade, M. J., N. A. Johnson, R. Jones, V. Siguel, and M. McNaughton. 1997. Genetic variation segregating in natural populations of Tribolium castaneum affecting traits observed in hybrids with T freemani. Genetics 147:1235-1247.
Wade, M. J., N. W. Chang, and M. McNaughton. 1995. Incipient speciation in the flour beetle, Tribolium confusum: premating isolation between natural populations. Heredity 75:453-459.