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Mitroshina, L. A. 1986. Specific composition and peculiarities of darkling beetles in south Ustyurt. Izvestiya Akademii Nauk Turkmenskoi SSR Seriya Biologicheskikh Nauk 4:76-78.
Mitroshina, L. A. 1988. On the fauna of tenebrionids (Coleoptera, Tenebrionidae) in the south Usturt. Zoologicheskii Zhurnal 67:1587-1589.
Mitroshina, L. A. 1990. Adaptive peculiarities of darkling beetles (Coleoptera, Tenebrionidae) to arid conditions of south Usturt. Ekologiya 1:121-128.