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Mordkovich, V. G. 1982. The role of the darkling beetles (Coleoptera, Tenebrionidae) in steppe ecosystems. Izvestiya Sibirskogo Otdeleniya Akademii Nauk SSSR Seriya Biologicheskikh Nauk 3:103-109.
Mordkovich, V. G. 2002. Species richness, population structure and functional significance of black-beetles (Coleoptera: Tenebrionidae) in steppes of Northern Asia. Russian Entomological Journal 11:57-68.
Mordkovich, V. G. 2006. Darkling beetles (Coleoptera: Tenebrionidae) in the steppe catenae. Trudy Russkogo Entomologicheskogo Obshchestva 77:235-240.
Mordkovich, V. G., and N. A. Afanas'ev. 1980. Transformation of steppe litter by darkling beetles (Coleoptera, Tenebrionidae. Ekologiya 3:56-62.
Mordkovich, V. G., and U. A. Maadyr Ool. 2001. Topo-ecological groups of darkling beetles (Coleoptera, Tenebrionidae) with consideration of its adaptive morphology and tropho-physiology. Russian Entomological Journal 10:213-218.
Mordkovich, V. G., and V. V. Volkovintser. 1973. Adaptive types of darkling-beetles (Coleoptera, Tenebrionidae) in natural condition gradient. Voprosy Ekologii Novosibirskii Gosudarstvennyi Universitet:2-2.
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Tshernyshev, S. E., and V. G. Mordkovich. 2002. New species of the genus Blaps Fabricius, 1775 (Coleoptera: Tenebrionidae) found in Siberia. Russian Entomological Journal 11:383-385.