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Colombini, I., L. Chelazzi, M. Fallaci, and L. Palesse. 1994. Zonation and surface activity of some tenebrionid beetles living on a mediterranean sandy beach. Journal of Arid Environments 28:215-230.
Fallaci, M., I. Colombini, L. Palesse, and L. Chelazzi. 1997. Spatial and temporal strategies in relation to environmental constraints of four tenebrionids inhabiting a Mediterranean coastal dune system. Journal of Arid Environments 37:45-64.