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Chen, G., K. Handel, and S. Roth. 2000. The maternal NF-[kappa]B/Dorsal gradient of Tribolium castaneum: dynamics of early dorsoventral patterning in a short-germ beetle. Development 127:5145-5156.
Handel, K., A. Basal, X. Fan, and S. Roth. 2005. Tribolium castaneum twist: gastrulation and mesoderm formation in a short-germ beetle. Development Genes and Evolution 215:13-31.
Handel, K., C. G. Grunfelder, S. Roth, and K. Sander. 2000. Tribolium embryogenesis: a SEM study of cell shapes and movements from blastoderm to serosal closure. Development Genes and Evolution 210:167-179.
Maxton Kuchenmeister, J., K. Handel, U. Schmidt Ott, S. Roth, and H. Jackle. 1999. Toll homolog expression in the beetle Tribolium suggests a different mode of dorsoventral patterning than in Drosophila embryos. Mechanisms of Development 83:107-114.
Roth, L. M. 1943. Studies on the gaseous secretions of Tribolium confusum DuVal. Annals of the Entomological Society of America 36:397-424.
van der Zee, M., N. Berns, and S. Roth. 2005. Distinct functions of the Tribolium zerknuellt genes in serosa specification and dorsal closure. Current Biology 15:624-636.