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Fattorini, S., P. Leo, and L. Salvati. 1999. Biogeographical observations on the darkling beetles of the Aegean Islands (Coleoptera, Tenebrionidae. Fragmenta Entomologica 31:339-375.
Fattorini, S., P. Leo, and L. Salvati. 2000. Levels of endemism in the Aegean tenebrionids (Coleoptera, Tenebrionidae. Biogeographia 21:429-440.