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Handel, K., C. G. Grunfelder, S. Roth, and K. Sander. 2000. Tribolium embryogenesis: a SEM study of cell shapes and movements from blastoderm to serosal closure. Development Genes and Evolution 210:167-179.
Schmidt Ott, U., K. Sander, and G. M. Technau. 1994. Expression of engrailed in embryos of a beetle and five dipteran species with special reference to the terminal regions. Roux's Archives of Developmental Biology 203:298-303.