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Enders, M. M., H. Schule, and J. R. Henschel. 1998. Sexual dimorphism, fighting success and mating tactics of male Onymacris plana Peringuey (Coleoptera: Tenebrionidae) in the Namib Desert. Ethology 104:1003-1019.
Enders, M., and H. Schule. 1993. Age identification of a Namib desert tenebrionid beetle. Journal of Arid Environments 24:211-214.