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Bedir, M. A., A. Shaban, F. M. Semida, and A. H. Abo Ghalia. 2002. Palatability of various plant leaves to three darkling beetle species (family: Tenebrionidae. Journal of the Egyptian German Society of Zoology 39:21-31.
Bedir, M. A., F. M. Semida, A. Shaban, and A. H. Abo Ghalia. 2005. Bioenergetic studies on three species of Egyptian tenebrionid beetles. Journal of the Egyptian German Society of Zoology 47:39-68.