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Stapp, P. 1997. Microhabitat use and community structure of darkling beetles (Coleoptera: Tenebrionidae) in shortgrass prairie: effects of season, shrub cover and soil type. American Midland Naturalist 137:298-311.
Stapp, P. 1997. Small mammal predation on darkling beetles (Coleoptera: Tenebrionidae) in pitfall traps. Southwestern Naturalist 42:352-355.