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Tsuda, Y., and T. Yoshida. 1984. Population biology of the broad-horned flour beetle, Gnathocerus cornutus (F) (Coleoptera: Tenebrionidae) 1 Life table and population parameters. Applied Entomology and Zoology 19:129-131.
Tsuda, Y., and T. Yoshida. 1985. Population biology of the broad-horned flour beetle, Gnathocerus cornutus (F) 2 Crowding effects of larvae on their survival and development. Researches on Population Ecology 27:77-85.