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Whitlock, M. 1993. Lack of correlation between heterozygosity and fitness in forked fungus beetles. Heredity 70:574-581.
Whitlock, M. C. 1992. Nonequilibrium population structure in forked fungus beetles: extinction, colonization, and the genetic variance among populations. American Naturalist 139:952-970.
Whitlock, M. C. 1994. Fission and the genetic variance among populations: the changing demography of forked fungus beetle populations. American Naturalist 143:820-829.