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Nakakita, H., O. Imura, and R. G. Winks. 1981. Hybridization between Tribolium freemani Hinton and Tribolium castaneum (Herbst), and some preliminary studies on the biology of Tribolium freemani (Coleoptera: Tenebrionidae. Applied Entomology and Zoology 16:209-215.