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de Almeida, M. C., A. A. Zacaro, and D. M. Cella. 2000. Cytogenetic analysis of Epicauta atomaria (Meloidae) and Palembus dermestoides (Tenebrionidae) with Xyp sex determination system using standard staining, C-bands, NOR and synaptonemal complex microspreading techniques. Hereditas 133:147-157.
Zacaro, A. A., M. C. de Almeida, and D. M. Cella. 2003. Recombination nodules in coleopteran species: Palembus dermestoides (Tenebrionidae) and Epicauta atomaria (Meloidae. Cytogenetic and Genome Research 103:185-191.