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Riddle, R. A., P. S. Dawson, and D. F. Zirkle. 1986. An experimental test of the relationship between genetic variation and environmental variation in Tribolium flour beetles. Genetics 112:391-404.
Zirkle, D. F., and P. S. Dawson. 1988. An experimental analysis of the relationships between fitness components and malic enzyme genotypes in Tribolium confusum. Biochemical Genetics 26:277-286.
Zirkle, D. F., and R. A. Riddle. 1983. Quantitative genetic response to environmental heterogeneity in Tribolium confusum. Evolution 37:637-639.
Zirkle, D. F., P. S. Dawson, and B. Lavie. 1988. An experimental analysis of the genetic relationships among life-history traits and emigration behavior in Tribolium castaneum. Oikos 53:391-397.