Aalbu, R. L., T. J. Spilman, and K. W. Brown. 1995. The systematic status of Amblycyphrus asperatus, Threnus niger, Pycnomorpha californica, Emmenastus rugosus, and Biomorphus tuberculatus Motschulsky (Coleoptera: Tenebrionidae). Proceedings of the Entomological Society of Washington 97:481-488.

Taxa (in this database) described in this work: