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De Los Santos, A., E. J. Alonso, E. Hernandez, and A. M. Perez. 2002. Environmental correlates of darkling beetle population size (Col Tenebrionidae) on the Canadas of Teide in Tenerife (Canary Islands. Journal of Arid Environments 50:287-308.
De Los Santos, A., L. A. Gomez Gonzalez, C. Alonso, C. D. Arbelo, and J. P. De Nicolas. 2000. Adaptive trends of darkling beetles (Col Tenebrionidae) on environmental gradients on the island of Tenerife (Canary Islands. Journal of Arid Environments 45:85-98.