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Ayal, Y., and O. Merkl. 1994. Spatial and temporal distribution of tenebrionid species (Coleoptera) in the Negev Highlands, Israel. Journal of Arid Environments 27:347-361.
Groner, E., and Y. Ayal. 2001. The interaction between bird predation and plant cover in determining habitat occupancy of darkling beetles. Oikos 93:22-31.
Krasnov, B., and Y. Ayal. 1995. Seasonal changes in darkling beetle communities (Coleoptera: Tenebrionidae) in the Ramon erosion cirque, Negev Highlands, Israel. Journal of Arid Environments 31:335-347.