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Carpaneto, G. M., and S. Fattorini. 2001. Spatial and seasonal organisation of a darkling beetle (Coleoptera, Tenebrionidae) community inhabiting a Mediterranean coastal dune system. Italian Journal of Zoology 68:207-214.
Carpaneto, G. M., and S. Fattorini. 2003. Seasonal occurrence and habitat distribution of tenebrionid beetles inhiabiting a Mediterranean coastal dune (Circeo National Park, Italy. Revue d'Ecologie la Terre et la Vie 58:293-306.
Fattorini, S., and G. M. Carpaneto. 2001. Tenebrionid density in Mediotyrrhenian coastal dunes: Habitat and seasonal variations (Coleoptera, Tenebrionidae. Fragmenta Entomologica 33:97-118.