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De Los Santos, A., F. Ferrer, J. P. De Nicolas, and T. O. Crist. 2006. Thermal habitat and life history of two congeneric species of darkling beetles (Coleoptera: Tenebrionidae) on Tenerife (Canary Islands. Journal of Arid Environments 65:363-385.
De Los Santos, A., J. P. De Nicolas, and F. Ferrer. 2002. Habitat selection and assemblage structure of darkling beetles (Col Tenebrionidae) along environmental gradients on the island of Tenerife (Canary Islands. Journal of Arid Environments 52:63-85.
De Los Santos, A., L. A. Gomez Gonzalez, C. Alonso, C. D. Arbelo, and J. P. De Nicolas. 2000. Adaptive trends of darkling beetles (Col Tenebrionidae) on environmental gradients on the island of Tenerife (Canary Islands. Journal of Arid Environments 45:85-98.