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Hanrahan, S. A., and W. H. Kirchner. 1994. Acoustic orientation and communication in desert tenebrionid beetles in sand dunes. Ethology 97:26-32.
Hanrahan, S. A., and W. H. Kirchner. 1997. The effect of wind on foraging activity of the tenebrionid beetle Lepidochora discoidalis in the sand dunes of the Namib Desert. South African Journal of Zoology 32:136-139.