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Bogachev, A. V., and O. L. Kryzhanovskii. 1955. A new species of Coleoptera, Tenebrionidae from western Turkmenistan. Entomologischeskoe Obozrenie 34:240-241.
Kryzhanovskii, O. L., and M. G. Nepesova. 1990. [Attempt at reconstruction of desert fauna origins in Turkmenistan - Tenebrionidae. Izvestiya Akademii Nauk Turkmenskoi SSR Seriya Biologicheskikh Nauk 4:121-128.