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Jockusch, E. L., T. A. Williams, and L. M. Nagy. 2004. The evolution of patterning of serially homologous appendages in insects. Development Genes and Evolution 214:324-338.
Nagy, L. M., and S. Carroll. 1994. Conservation of wingless patterning functions in the short-germ embryos of Tribolium castaneum. Nature 367:460-463.