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Brown, S. J., J. K. Parrish, R. E. Denell, and R. W. Beeman. 1994. Genetic control of early embryogenesis in the red flour beetle, Tribolium castaneum. American Zoologist 34:343-352.
Brown, S. J., J. K. Parrish, R. W. Beeman, and R. E. Denell. 1997. Molecular characterization and embryonic expression of the even-skipped ortholog of Tribolium castaneum. Mechanisms of Development 61:165-173.