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Gomez Ruano, R., and L. Silvela. 1998. Total weight loss (TWL) An easy to score, fitness related, heritable trait in Tribolium castaneum Herbst. Tribolium Information Bulletin 38:259-260.
Ruano, R. G., L. S. Silvela, C. Lopez Fanjul, and M. A. Toro. 1996. Changes in the additive variance of a fitness-related trait with inbreeding in Tribolium castaneum. Journal of Animal Breeding and Genetics 113:93-97.
Silvela, L., I. de la Pena, and R. Gomez Ruano. 1999. Selection under negative linkage disequilibrium Random mating versus inbreeding. Heredity 82:598-604.