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Tanner, V. M. 1927. A preliminary study of the genitalia of female Coleoptera. Transactions of the American Entomological Society 53:5-50.
Tanner, V. M. 1961. A Checklist of the species of Eleodes and descriptions of new species (Coleoptera-Tenebrionidae). Great Basin Naturalist 21:55-78.
Tanner, V. M., and W. A. Packham. 1965. Tenebrionid beetles of the Nevada Test Site. Brigham Young University Science Bulletin 4:1-44.