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Harris, A. C., and I. L. Weatherall. 1990. Objective evaluation of colour variation in the sand-burrowing beetle Chaerodes trachyscelides White (Coleoptera: Tenebrionidae) by instrumental determination of CIELAB values. Journal of the Royal Society of New Zealand 20:253-259.
Harris, A. C., and I. L. Weatherall. 1991. Geographic variation for colour in the sand-burrowing beetle Chaerodes trachyscelides White (Coleoptera: Tenebrionidae) on New Zealand beaches analysed using CIELAB L values. Biological Journal of the Linnean Society 44:93-104.