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Ferrer, J. 1999. Contribution a la connaissance du genre Phrynocolus Lacordaire, 1859 Designation du neotype de Phrynocolus humilis Wilke, 1921 (Coleoptera, Tenebrionidae) Contribution to the knowledge of the genus Phrynocolus Lacordaire, 1859 Designation of the neotype of Phrynocolus humilis Wilke, 1921 (Coleoptera, Tenebrionidae. Nouvelle Revue d'Entomologie 16:23-26.

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