Ferrer, J., and F. Ødegaard. 2005. New species of darkling beetles from Central America with systematic notes (Coleoptera: Tenebrionidae. Annales Zoologici 55:633-661.
Ferrer, J., and F. Ødegaard. 2005. New species of darkling beetles from Central America with systematic notes (Coleoptera: Tenebrionidae. Annales Zoologici 55:633-661.