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Medvedev, G. S. 1968. Coleoptera Darkling beetles (Tenebrionidae) Subfamily Opatrinae Tribes Platynotini, Dendarini, Pedinini, Dissonomini, Pachypterini, Opatrini (part) and Heterotarsini Vol XIX Part 2 Fauna USSR NS No 97. Nauka, Leningrad.

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