Penrith, M. L. 1981. Revision of the Zophosini (Coleoptera: Tenebrionidae) Part 4 Twelve subgenera from arid southern Africa. Cimbebasia 6:125-164.
Penrith, M. L. 1981. Revision of the Zophosini (Coleoptera: Tenebrionidae) Part 4 Twelve subgenera from arid southern Africa. Cimbebasia 6:125-164.