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Penrith, M. L. 1983. Revision of the Zophosini (Coleoptera: Tenebrionidae) Part 8 The Palaearctic species group of the subgenus Oculosis Penrith, the subgenus Cheirosis Deyrolle, and a monotypical subgenus from Socotra. Cimbebasia 6:369-384.

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